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Different stages are used for different stages of the baby's development. The hole in the nipple is actually larger, or in some there are more holes which allows a faster flow. if you look at the different stages it will tell you on the package the age the suggest the stage be used for. Also if your baby seems to be sucking too hard and getting nothing or even like he/she seems to be drowning because of too much, you would want to change nipples.

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16y ago
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Q: What do the levels on baby bottle nipples mean?
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