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I Believe she's a Beechcraft C-45 Expeditor that's Seen Better days lol. But The C-45 was the World War II military version of the popular Beechcraft Model 18 commercial light transport. Beech built a total of 4,526 of these aircraft for the Army Air Forces between 1939 and 1945 in four versions, the AT-7 Navigator navigation trainer, the AT - 1 Kansan bombing-gunnery trainer, the C-45 Expeditor utility transport and the F-2 for aerial Photography and mapping. The AT-7 and AT-11 versions were well-known to WWII navigators and bombardiers, for most of these men received their training in these aircraft. Thousands of AAF pilot cadets also were given advanced training in twin-engine Beech airplanes.

During the early 1950s, Beech completely rebuilt 900 C-45s for the Air Force. They received new serial numbers and were designated C-45Gs and C-45Hs, remaining in service until 1963 for administrative and light cargo duties. or for fairing lions, hippos, zebras,giraffes,lemurs monkeys and penguins around I'm sure lol

[addendum]

While the outside looks like a Beech C-45 / D-18 (I seem to recall passenger remains appeared to be in civilian attire) or even a Lockheed Electra the INTERIOR looks too big for such a size aircraft, appear to be more the size of a DC-3 inside the passenger compartment. Meanwhile the flight deck looks like neither.

This leads me to believe that while the were inspired by the aforementioned aircraft, the airplane in the movie is an entirely fictional construct of the animators/modelers imaginations.

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Q: What is the plane in Madagascar 2?
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